List of vulnerable reptiles
- Extinct in the wild (EW): 2 species
- Critically endangered (CR): 196 species
- Endangered (EN): 382 species
- Vulnerable (VU): 411 species
- Near threatened (NT, LR/cd): 329 species
- Least concern (LC): 2,900 species
- Data deficient (DD): 910 species
As of September 2016, the International Union for Conservation of Nature (IUCN) lists 411 vulnerable reptile species.[1] 8.0% of all evaluated reptile species are listed as vulnerable. The IUCN also lists ten reptile subspecies as vulnerable.
No subpopulations of reptiles have been evaluated as vulnerable by the IUCN.
For a species to be assessed as vulnerable to extinction the best available evidence must meet quantitative criteria set by the IUCN designed to reflect "a high risk of extinction in the wild". Endangered and critically endangered species also meet the quantitative criteria of vulnerable species, and are listed separately. See: List of endangered reptiles, List of critically endangered reptiles. Vulnerable, endangered and critically endangered species are collectively referred to as threatened species by the IUCN.
Additionally 910 reptile species (18% of those evaluated) are listed as data deficient, meaning there is insufficient information for a full assessment of conservation status. As these species typically have small distributions and/or populations, they are intrinsically likely to be threatened, according to the IUCN.[2] While the category of data deficient indicates that no assessment of extinction risk has been made for the taxa, the IUCN notes that it may be appropriate to give them "the same degree of attention as threatened taxa, at least until their status can be assessed."[3]
This is a complete list of vulnerable reptile species and subspecies evaluated by the IUCN.
Turtles and tortoises
There are 62 turtle species assessed as vulnerable.
Tortoises
- African spurred tortoise (Centrochelys sulcata)
- Volcan wolf giant tortoise (Chelonoidis becki)
- San Cristobal giant tortoise (Chelonoidis chathamensis)
- Chaco tortoise (Chelonoidis chilensis)
- Yellow-footed tortoise (Chelonoidis denticulata)
- Indian star tortoise (Geochelone elegans)
- Aldabra giant tortoise (Geochelone gigantea)
- Desert tortoise (Gopherus agassizii)
- Bolson tortoise (Gopherus flavomarginatus)
- Gopher tortoise (Gopherus polyphemus)
- Nama padloper (Homopus solus)
- Travancore tortoise (Indotestudo travancorica)
- Home's hinge-back tortoise (Kinixys homeana)
- Pancake tortoise (Malacochersus tornieri)
- Impressed tortoise (Manouria impressa)
- Spur-thighed tortoise (Testudo graeca)
- Russian tortoise (Testudo horsfieldii)
Geoemydids
- Amboina box turtle (Cuora amboinensis)
- Black pond turtle (Geoclemys hamiltonii)
- Brahminy river turtle (Hardella thurjii)
- Giant Asian pond turtle (Heosemys grandis)
- Mekong snail-eating turtle (Malayemys subtrijuga)
- Tricarinate hill turtle (Melanochelys tricarinata)
- Burmese eyed turtle (Morenia ocellata)
- Indian eyed turtle (Morenia petersi)
- Malayan flat-shelled turtle (Notochelys platynota)
- Black marsh turtle (Siebenrockiella crassicollis)
Trionychids
- Asiatic softshell turtle (Amyda cartilaginea)
- Senegal flapshell turtle (Cyclanorbis senegalensis)
- Indian softshell turtle (Nilssonia gangetica)
- Indian peacock softshell turtle (Nilssonia hurum)
- Leith's softshell turtle (Nilssonia leithii)
- New Guinea giant softshell turtle (Pelochelys bibroni)
- Chinese softshell turtle (Pelodiscus sinensis)
Chelids
- Parker's snake-necked turtle (Chelodina parkeri)
- White-bellied snapping turtle (Elseya branderhorsti)
- Brazilian snake-necked turtle (Hydromedusa maximiliani)
- Zulia toad-headed sideneck (Mesoclemmys zuliae)
- Fitzroy River turtle (Rheodytes leukops)
Emydids
- Western pond turtle (Actinemys marmorata)
- Barbour's map turtle (Graptemys barbouri)
- Yellow-blotched map turtle (Graptemys flavimaculata)
- Ringed map turtle (Graptemys oculifera)
- Common box turtle (Terrapene carolina)
- Hispaniolan slider (Trachemys decorata)
- Big Bend slider (Trachemys gaigeae)
- Ornate slider (Trachemys ornata)
- Jamaican slider (Trachemys terrapen)
- Yaqui slider (Trachemys yaquia)
Other turtle species
- Loggerhead sea turtle (Caretta caretta)
- Pig-nosed turtle (Carettochelys insculpta)
- Central American snapping turtle (Chelydra rossignonii)
- Leatherback sea turtle (Dermochelys coriacea)
- Central American mud turtle (Kinosternon angustipons)
- Dunn's mud turtle (Kinosternon dunni)
- Olive ridley sea turtle (Lepidochelys olivacea)
- Alligator snapping turtle (Macrochelys temminckii)
- Big-headed Amazon River turtle (Peltocephalus dumerilianus)
- Turkana mud turtle (Pelusios broadleyi)
- Red-headed Amazon River turtle (Podocnemis erythrocephala)
- Six-tubercled Amazon River turtle (Podocnemis sextuberculata)
- Yellow-spotted river turtle (Podocnemis unifilis)
Crocodilia species
- American crocodile (Crocodylus acutus)
- Mugger crocodile (Crocodylus palustris)
- Dwarf crocodile (Osteolaemus tetraspis)
- False gharial (Tomistoma schlegelii)
Tuatara
- Brothers Island tuatara (Sphenodon guntheri)
Lizards
There are 244 species and nine subspecies of lizard assessed as vulnerable.
Iguanids
Includes iguanas and related species.
Species
- Marine iguana (Amblyrhynchus cristatus)
- Santa Fe land iguana (Conolophus pallidus)
- Galapagos land iguana (Conolophus subcristatus)
- Balsas spiny-tailed iguana (Ctenosaura clarki)
- Yucatán spiny-tailed iguana (Ctenosaura defensor)
- Nolasco spiny-tailed iguana (Ctenosaura nolascensis)
- Rhinoceros iguana (Cyclura cornuta)
- Northern Bahamian rock iguana (Cyclura cychlura)
- Clouded rock iguana (Cyclura nubila)
Subspecies
- Isabella marine iguana (Amblyrhynchus cristatus albemarlensis)
- Amblyrhynchus cristatus cristatus
- Santa Cruz marine iguana (Amblyrhynchus cristatus hassi)
- Pinta marine iguana (Amblyrhynchus cristatus sielmanni)
- Española marine iguana (Amblyrhynchus cristatus venustissimus)
- Cuban rock iguana (Cyclura nubila nubila)
Flap-footed lizards
- Hermite Island worm-lizard (Aprasia rostrata)
- Striped legless lizard (Delma impar)
- Striped-tailed delma (Delma labialis)
- Collared delma (Delma torquata)
- Bronzeback snake-lizard (Ophidiocephalus taeniatus)
- Brigalow scaly-foot (Paradelma orientalis)
Anguids
Includes slowworms, glass lizards, and alligator lizards.
- Anzuetoi arboreal alligator lizard (Abronia anzuetoi)
- Mixtecan arboreal alligator lizard (Abronia mixteca)
- Oaxaca arboreal alligator lizard (Abronia oaxacae)
- Bromeliad arboreal alligator lizard (Abronia taeniata)
- Bocourt's arboreal alligator lizard (Abronia vasconcelosii)
- Hispaniolan khaki galliwasp (Celestus curtissi)
- Panamint alligator lizard (Elgaria panamintina)
- Ophisaurus hainanensis
Girdled lizards
- Giant girdled lizard (Cordylus giganteus)
- McLachlan's girdled lizard (Cordylus mclachlani)
- Dwarf crag lizard (Cordylus nebulosus)
- Tasman's girdled lizard (Cordylus tasmani)
- Armadillo girdled lizard (Ouroborus cataphractus)
- Emperor flat lizard (Platysaurus imperator)
Chameleons
- Northern leaf chameleon (Brookesia ebenaui)
- Mount d'Ambre leaf chameleon (Brookesia tuberculata)
- Brookesia vadoni
- Calumma capuroni
- Hooded chameleon (Calumma cucullatum)
- Marojejy peak chameleon (Calumma jejy)
- O'Shaughnessy's chameleon (Calumma oshaughnessyi)
- Calumma peyrierasi
- Tsaratanana chameleon (Calumma tsaratananense)
- Blunt-nosed chameleon (Calumma tsycorne)
- Antimena chameleon (Furcifer antimena)
- Jewelled chameleon (Furcifer campani)
- Labord's chameleon (Furcifer labordi)
- Petter's chameleon (Furcifer petteri)
- Rhinoceros chameleon (Furcifer rhinoceratus)
- Mount Kenya sentinel chameleon (Kinyongia excubitor)
- Elongate leaf chameleon (Palleon nasus)
- Mahenge pygmy chameleon (Rhampholeon beraduccii)
- Marshall's pygmy chameleon (Rhampholeon marshalli)
- Mount Chiperone pygmy chameleon (Rhampholeon nebulauctor)
- Four-horned chameleon (Trioceros quadricornis)
- Trioceros serratus
Plated lizards
- Breyer's long-tailed seps (Tetradactylus breyeri)
- Tracheloptychus petersi
- Zonosaurus anelanelany
- Zonosaurus boettgeri
- Zonosaurus maximus
- Four-lined girdled lizard (Zonosaurus quadrilineatus)
Anoles
- Anolis barkeri
- Navassa anole (Anolis longiceps)
- Rueda's anole (Anolis maculigula)
- Anolis muralla
- Anolis naufragus
- Crab cay anole (Anolis pinchoti)
- Bearded anole (Anolis pogus)
Gekkonids
- Giant bronze gecko (Ailuronyx trachygaster)
- Blaesodactylus boivini
- Christinus guentheri
- Mt Elgon forest gecko (Cnemaspis elgonensis)
- Indian day gecko (Cnemaspis indica)
- Das's day gecko (Cnemaspis indraneildasii)
- Jerdon's day gecko (Cnemaspis jerdonii)
- Vellore day gecko (Cnemaspis otai)
- Niah cave gecko (Cyrtodactylus cavernicolus)
- Chiku bent-toed gecko (Cyrtodactylus hidupselamanya)
- Gekko ernstkelleri
- Gigante narrow-disked gecko (Gekko gigante)
- Peking gecko (Gekko swinhonis)
- White-striped viper gecko (Hemidactylus albofasciatus)
- Hemidactylus gujaratensis
- Satara gecko (Hemidactylus sataraensis)
- Rotuma forest gecko (Lepidodactylus gardineri)
- Lepidodactylus listeri
- Viti forest gecko (Lepidodactylus manni)
- Tiny scaled gecko (Lygodactylus bivittis)
- Blanc's dwarf gecko (Lygodactylus blanci)
- Usambara dwarf gecko (Lygodactylus gravis)
- Madagascar dwarf gecko (Lygodactylus madagascariensis)
- Methuen's dwarf gecko (Lygodactylus methueni)
- Matoatoa brevipes
- Lesser night gecko (Nactus coindemirensis)
- Serpent Island gecko (Nactus serpensinsula)
- Angel's petite gecko (Paragehyra petiti)
- Grandidier's Madagascar ground gecko (Paroedura androyensis)
- Paroedura vazimba
- Phelsuma breviceps
- Phelsuma hielscheri
- Island day gecko (Phelsuma nigristriata)
- Standing's day gecko (Phelsuma standingi)
- Orange-spotted smooth-scaled gecko (Pseudogekko brevipes)
- Uroplatus ebenaui
- Uroplatus giganteus
- Henkel's leaf-tailed gecko (Uroplatus henkeli)
- Uroplatus malama
Wall lizards
Species
- Acanthodactylus felicis
- Leopard fringe-fingered lizard (Acanthodactylus pardalis)
- Darevskia alpina
- Mosor rock lizard (Dinarolacerta mosorensis)
- Iberian rock lizard (Iberolacerta monticola)
- Skyros wall lizard (Podarcis gaigeae)
- Podarcis levendis
- Milos wall lizard (Podarcis milensis)
Subspecies
- Zootoca vivipara pannonica
Skinks
Species
- Woodbush legless skink (Acontias rieppeli)
- Guinea lidless skink (Afroablepharus africana)
- Amphiglossus alluaudi
- Amphiglossus anosyensis
- Yellow skink (Amphiglossus ardouini)
- Amphiglossus mandokava
- Splendid skink (Amphiglossus splendidus)
- Long-legged worm-skink (Anomalopus mackayi)
- Mandjélia litter skink (Caledoniscincus terma)
- Günther's cylindrical skink (Chalcides guentheri)
- Manuel's skink (Chalcides manueli)
- Small three-toed skink (Chalcides minutus)
- Glorioso snake eyed skink (Cryptoblepharus gloriosus)
- Lancelin Island skink (Ctenotus lancelini)
- Hamelin ctenotus (Ctenotus zastictus)
- Griffin's keel-scaled tree skink (Dasia griffini)
- Erronan treeskink (Emoia erronan)
- Loyalty Islands emoia (Emoia loyaltiensis)
- Viti copper-headed skink (Emoia parkeri)
- Emoia tuitarere
- Gracile burrowing skink (Graciliscincus shonae)
- Kaestlea laterimaculata
- New Caledonian leopard skink (Lacertoides pardalis)
- Telfair's skink (Leiolopisma telfairii)
- Ruwenzori four toed skink (Leptosiaphos meleagris)
- Great desert skink (Liopholis kintorei)
- Ashwamedh writhing skink (Lygosoma ashwamedhi)
- Madascincus nanus
- Marmorosphax boulinda
- Montane marble-throated skink (Marmorosphax montana)
- Gracile dwarf skink (Nannoscincus gracilis)
- Earless dwarf skink (Nannoscincus mariei)
- Rankin's dwarf skink (Nannoscincus rankini)
- Florida sand skink (Neoseps reynoldsi)
- Pedra Branca skink (Niveoscincus palfreymani)
- Robust skink (Oligosoma alani)
- Falla's skink (Oligosoma fallai)
- Grand skink (Oligosoma grande)
- Chevron Skink (Oligosoma homalonotum)
- Lord Howe Island skink (Oligosoma lichenigera)
- Macgregor's skink (Oligosoma macgregori)
- Small-scaled skink (Oligosoma microlepis)
- New Zealand striped skink (Oligosoma striatum)
- Scree skink (Oligosoma waimatense)
- Whitaker's skink (Oligosoma whitakeri)
- Paracontias kankana
- Parvoscincus sisoni
- Duges' skink (Plestiodon dugesii)
- Pseudoacontias unicolor
- Günther's dwarf burrowing skink (Scelotes guentheri)
- Kasner's dwarf burrowing skink (Scelotes kasneri)
- Red-tailed shiny skink (Sigaloseps ruficauda)
- Orange-bellied burrowing skink (Simiscincus aurantiacus)
- Sphenomorphus knollmanae
- Trachylepis dumasi
- Trachylepis lavarambo
- Trachylepis loluiensis
- Trachylepis tavaratra
- Wright's skink (Trachylepis wrightii)
- Aubrey's whiptailed skink (Tropidoscincus aubrianus)
- Lomi's blind legless skink (Typhlosaurus lomiae)
Subspecies
- Egernia stokesii aethiops
- Jurien Bay rock-skink (Liopholis pulchra longicauda)
Spectacled lizards
- Ruthven's anadia (Anadia pulchella)
- Bresslau's bachia (Bachia bresslaui)
- Psilophthalmus paeminosus
- Werner's largescale lizard (Ptychoglossus bicolor)
- Shiny lightbulb lizard (Riama laevis)
- Riama stigmatoral
- Key tegu (Teuchocercus keyi)
Teiids
Includes whiptails and tegus.
- Censky's ameiva (Ameiva corax)
- Sombrero ameiva (Ameiva corvina)
- Inagua ameiva (Ameiva maynardii)
- Aspidoscelis catalinensis
- Little white whiptail (Aspidoscelis gypsi)
- Aspidoscelis labialis
- San Pedro Martir whiptail (Aspidoscelis martyris)
- Saint Lucia whiptail (Cnemidophorus vanzoi)
Dragon lizards
- Montane rock agama (Agama montana)
- Gunther's bloodsucker (Bronchocela smaragdina)
- Rough-nosed horned lizard (Ceratophora aspera)
- Yinnietharra rock dragon (Ctenophorus yinnietharra)
- Draco mindanensis
- Philippine sailfin lizard (Hydrosaurus pustulatus)
- Persian toad agame (Phrynocephalus persicus)
- Strauch's toad agama (Phrynocephalus strauchi)
- Savigny's agama (Trapelus savignii)
- Grassland earless dragon (Tympanocryptis pinguicolla)
- Egyptian mastigure (Uromastyx aegyptia)
- Omani spiny-tailed lizard (Uromastyx thomasi)
Phyllodactylids
- Asaccus montanus
- Belize leaf-toed gecko (Phyllodactylus insularis)
- Western leaf-toed gecko (Phyllodactylus lepidopygus)
- Boavista wall gecko (Tarentola boavistensis)
- Helmeted gecko (Tarentola chazaliae)
Phrynosomatids
- Dunes sagebrush lizard (Sceloporus arenicolus)
- Sceloporus maculosus
- Sceloporus megalepidurus
- Sceloporus oberon
- Urosaurus clarionensis
- Uta encantadae
- Uta lowei
- Uta palmeri
- Uta tumidarostra
Liolaemids
- Liolaemus dicktracyi
- Liolaemus famatinae
- Hellmich's tree iguana (Liolaemus hellmichi)
- Liolaemus lutzae
- Liolaemus morenoi
- Liolaemus occipitalis
- Liolaemus tandiliensis
- Thorntail mountain lizard (Phymaturus mallimaccii)
Other lizard species
- Wiegmann's tree lizard (Anisolepis undulatus)
- Reticulated collared lizard (Crotaphytus reticulatus)
- Los archipelago worm lizard (Cynisca leonina)
- Cynisca nigeriensis
- Red-eyed woodlizard (Enyalioides oshaughnessyi)
- Stephen's Island gecko (Hoplodactylus stephensi)
- East Plana curlytail (Leiocephalus greenwayi)
- Gaige's tropical night lizard (Lepidophyma gaigeae)
- Cave tropical night lizard (Lepidophyma micropholis)
- Ringed spinytail iguana (Morunasaurus annularis)
- Bavay's giant gecko (Rhacodactylus chahoua)
- Crested gecko (Rhacodactylus ciliatus)
- Sarasin's giant gecko (Rhacodactylus sarasinorum)
- Banded-toed gecko (Saurodactylus fasciatus)
- Rough-banded sphaero (Sphaerodactylus callocricus)
- Grenadines sphaero (Sphaerodactylus kirbyi)
- Cuban broad-banded sphaero (Sphaerodactylus torrei)
- Peracca's whorltail iguana (Stenocercus festae)
- Stenocercus torquatus
- Komodo dragon (Varanus komodoensis)
- Gray's monitor (Varanus olivaceus)
- Sandstone night lizard (Xantusia gracilis)
- Xenosaurus grandis
Snakes
There are 100 species and one subspecies of snake assessed as vulnerable.
Pseudoxyrhophiids
- Brygophis coulangesi
- Uluguru forest snake (Buhoma procterae)
- Compsophis zeny
- Liophidium therezieni
- Grandidier's water snake (Liopholidophis grandidieri)
- Lycodryas citrinus
- Lycodryas inornatus
- Pararhadinaea melanogaster
- Pseudoxyrhopus oblectator
- Pseudoxyrhopus sokosoko
- Yellow-striped water snake (Thamnosophis stumpffi)
Vipers
Species
- Barbour's short-headed viper (Atheris barbouri)
- Usambara eyelash viper (Atheris ceratophora)
- Plain mountain adder (Bitis inornata)
- Namaqua dwarf adder (Bitis schneideri)
- Yellow-blotched palm pit viper (Bothriechis aurifer)
- Rowley's palm pit viper (Bothriechis rowleyi)
- Piraja's lancehead (Bothrops pirajai)
- Crotalus stejnegeri
- Hon Son pit viper (Cryptelytrops honsonensis)
- Ruby-eyed green pitviper (Cryptelytrops rubeus)
- Shedao Island pit viper (Gloydius shedaoensis)
- Montivipera albicornuta
- Ophryacus undulatus
- Cameron highlands pit viper (Popeia nebularis)
- Caucasus subalpine viper (Vipera dinniki)
- Iranian mountain steppe viper (Vipera ebneri)
- Alburzi viper (Vipera eriwanensis)
- Lataste's viper (Vipera latastei)
- Meadow viper (Vipera ursinii)
Subspecies
- Atlantic bushmaster (Lachesis muta rhombeata)
Dipsadids
- Stuart's burrowing snake (Adelphicos veraepacis)
- Modest ground snake (Atractus modestus)
- Northern ground snake (Atractus nicefori)
- Roule's ground snake (Atractus roulei)
- Tropical forest snake (Calamodontophis paucidens)
- Peters' running snake (Coniophanes dromiciformis)
- Dixon's ground snake (Erythrolamprus atraventer)
- Geophis juliai
- Geophis nephodrymus
- Southern hognose snake (Heterodon simus)
- Omoadiphas aurula
- Philodryas livida
- Ribbon graceful brown snake (Rhadinaea fulvivittis)
- Monte Cristi graceful brown snake (Rhadinella montecristi)
- Rhadinella pegosalyta
- Tantalophis discolor
Elapids
- Pygmy copperhead (Austrelaps labialis)
- South Andaman krait (Bungarus andamanensis)
- Red River krait (Bungarus slowinskii)
- Ornamental snake (Denisonia maculata)
- Lake Cronin snake (Echiopsis atriceps)
- Dunmall's snake (Furina dunmalli)
- Broad-headed snake (Hoplocephalus bungaroides)
- Lake Taal snake (Hydrophis semperi)
- Crocker's sea snake (Laticauda crockeri)
- Katuali (Laticauda schistorhynchus)
- Solomons black-banded krait (Loveridgelaps elapoides)
- Oaxacan coral snake (Micrurus ephippifer)
- Chinese cobra (Naja atra)
- Mandalay spitting cobra (Naja mandalayensis)
- Indochinese spitting cobra (Naja siamensis)
- King cobra (Ophiophagus hannah)
- Woodlark Island forest snake (Toxicocalamus longissimus)
Colubrids
- Dryocalamus philippinus
- Philippine dryophiops (Dryophiops philippina)
- Cloud forest parrot snake (Leptophis modestus)
- Rendahl's wolf snake (Lycodon paucifasciatus)
- Short-tailed kukri snake (Oligodon brevicauda)
- Walnut kukri snake (Oligodon juglandifer)
- Lacroix kukri snake (Oligodon lacroixi)
- Spotted-bellied short-headed snake (Oligodon modestum)
- Oligodon pulcherrimus
- Moellendorff's trinket snake (Orthriophis moellendorfi)
- Tantilla boipiranga
- Jan's centipede snake (Tantilla jani)
- Tantilla psittaca
- Fruhstorfer's mountain snake (Tetralepis fruhstorferi)
- Usambara vine snake (Thelotornis usambaricus)
Keelbacks
- Adelophis copei
- Maki's keelback (Hebius miyajimae)
- Large-headed water snake (Natrix megalocephala)
- Storeria hidalgoensis
- Giant garter snake (Thamnophis gigas)
- Thamnophis scaliger
- Tropidonophis negrosensis
Other snake species
- Hainan odd-scaled snake (Achalinus hainanensis)
- Amami Takachiho snake (Achalinus werneri)
- Stadelman's worm snake (Amerotyphlops stadelmani)
- Amerotyphlops tycherus
- Jamaican boa (Chilabothrus subflavus)
- Longhead water snake (Enhydris longicauda)
- Exiliboa placata
- Dwarf wolf snake (Lycophidion nanus)
- Two-lined black earth snake (Melanophidium bilineatum)
- Mcnamara's burrowing snake (Pseudorabdion mcnamarae)
- Pseudorabdion talonuran
- Burmese python (Python bivittatus)
- Python kyaiktiyo
- Christmas Island blind snake (Ramphotyphlops exocoeti)
- Phipson's earth snake (Uropeltis phipsonii)
See also
- Lists of IUCN Red List vulnerable species
- List of least concern reptiles
- List of near threatened reptiles
- List of endangered reptiles
- List of critically endangered reptiles
- List of recently extinct reptiles
- List of data deficient reptiles
References
- ↑ "IUCN Red List version 2016-2". The IUCN Red List of Threatened Species. International Union for Conservation of Nature and Natural Resources (IUCN). Retrieved 8 September 2016.
- ↑ "Limitations of the Data". The IUCN Red List of Threatened Species. Union for Conservation of Nature and Natural Resources (IUCN). Retrieved 11 January 2016.
- ↑ "2001 Categories & Criteria (version 3.1)". The IUCN Red List of Threatened Species. Union for Conservation of Nature and Natural Resources (IUCN). Retrieved 11 January 2016.